It seems there are (at least) doubled quads in the hole-areas. (And ANIMATOR can’t this brics display correctly while WINGRAF shows that there is something like brics and also shows, that there are quad-problems.)
For better debugging you can extrude only a part:
“tran quad prop x001 dz 11.7 xmax -29.39” instead “EXTR TYP VON BIS PATH V1 V2 V3 DIV …”
And you can pick quadnumbers in ANIMATOR and delete this Quads in a predefined MSHA (before! the running of the extruding MSHA).
If you delete a number of the in ANIMAOTOR as extruded shown quads: there will be a hole in the quad- and bric-structure.
If you delete a other quad of the extruding area: there is no whole, becouse there is yet another quad.